Type: COLOMBIA: BOYACA: subparamo on dry rocky slope in thicket of Arcytophyllum nitidum and Gaylussacia buxifolia associated with Espeletia glandulosa, Paepalanthus and Cladonia spp., Paramo de Guina, Santa Rosita, 3300 m alt, 5 May 1973, Cleef 9740 (COL, holotype; U).