Type: J. Cuatrecasas & T. R. Soderstrom 27308 (holotype US), Massif of Bogota, eastern slope, drainage of Rio Negro, knife crests between Fomeque and Laguna de Chingaza, between El Paval and Cuchillero, 900-1100 m alt, Cundinamarca, Colombia, 7 Feb 1969.