Type: MÉXICO. Oaxaca. 39.6 mi N of MEX 195 E of the city of Oaxaca along MEX 175 to Ixtlán de Juarez and Tuxtepec, 5600 ft, 28 Jul 1987, J. Utley & K. Utley 7961 (holotype: MEXU; isotypes: B, C, CAS, GH, M, MICH, MO, NY, US, USF, XAL). Figures 1, 2, and 3.